true blood-sucking bats or vampyres (Phyllostomidae), the capybara, the largest of the rodents, the carnivorous coatimondi (Nasua), and a great many other forms, are also exclusively characteristic of South America.
"In Peru and Chili," says Humboldt, "the region of the grasses, which is at an elevation of from 12,300 to 15,400 feet, is inhabited by crowds of lama, guanaco, and alpaca. These quadrupeds, which here represent the genus camel of the ancient continent, have not extended themselves either to Brazil or Mexico; because, during their journey, they must necessarily have descended into regions that were too hot for them."[876] In this passage it will be seen that the doctrine of "specific centres" is tacitly assumed.
Quadrupeds in Islands.--Islands remote from continents, especially those of small size, are either destitute of quadrupeds, except such as have been conveyed to them by man, or contain species peculiar to them. In the Galapagos archipelago no indigenous quadrupeds were found except one mouse, which is supposed to be distinct from any hitherto found elsewhere. A peculiar species of fox is indigenous in the Falkland Islands, and a rat in New Zealand, which last country, notwithstanding its magnitude, is destitute of other mammalia, except bats, and these, says Dr. Prichard, may have made their way along the chain of islands which extend from the shores of New Guinea far into the Southern Pacific. The same author remarks, that among the various groups of fertile islands in the Pacific, no quadrupeds have been met with except the rat and a few bats as above mentioned, and the dog and hog, which appear to have been conveyed thither by the natives from New Guinea. "Rats are to be found even on some desert islands, whither they may have been conveyed by canoes which have occasionally approached the shore. It is known, also, that rats occasionally swim in large numbers to considerable distances."[877]
Geographical range of the Cetacea.--It is natural to suppose that the geographical range of the different species of Cetacea should be less correctly ascertained than that of the terrestrial mammifers. It is, however, well known that the whales which are obtained by our fishers in the South Seas are distinct from those of the North; and the same dissimilarity has been found in all the other marine animals, of the same class, so far as they have yet been studied by naturalists.
Dispersion of quadrupeds.--Let us now inquire what facilities the various land quadrupeds enjoy of spreading themselves over the surface of the earth. In the first place, as their numbers multiply, all of them, whether they feed on plants, or prey on other animals, are disposed to scatter themselves gradually over as wide an area as is accessible to them. But before they have extended their migrations over a large space, they are usually arrested either by the sea, or a zone of uncongenial climate, or some lofty and unbroken chain of mountains, or a tract already occupied by a hostile and more powerful species.