HKB
to such definitions, unless we have previously made up our minds on some of the most momentous of all the enigmas with which the human intellect ever attempted to grapple.
It is now thirty years since I gave an analysis in the first edition of my "Principles of Geology" (volume 2 1832) of the views which had been put forth by Lamarck, in the beginning of the century, on this subject. In that interval the progress made in zoology and botany, both in augmenting the number of known animals and plants, and in studying their physiology and geographical distribution and above all in examining and describing fossil species, is so vast that the additions made to our knowledge probably exceed all that was previously known; and what Lamarck then foretold has come to pass; the more new forms have been multiplied, the less are we able to decide what we mean by a variety, and what by a species. In fact, zoologists and botanists are not only more at a loss than ever how to define a species, but even to determine whether it has any real existence in nature, or is a mere abstraction of the human intellect, some contending that it is constant within certain narrow and impassable limits of variability, others that it is capable of indefinite and endless modification.
Before I attempt to explain a great step, which has recently been made by Mr. Darwin and his fellow-labourers in this field of inquiry, I think it useful to recapitulate in this place some of the leading features of Lamarck's system, without attempting to adjust the claims of some of his contemporaries (Geoffroy St. Hilaire in particular) to share in the credit of some of his original speculations.
From the time of Linnaeus to the commencement of the present century, it seemed a sufficient definition of the term species to say that "a species consisted of individuals all resembling each other, and reproducing their like by generation." But Lamarck after having first studied botany with success, had then turned his attention to conchology, and soon became aware that in the newer (or Tertiary) strata of the earth's crust there were a multitude of fossil species of shells, some of them identical with living ones, others simply varieties of the living, and which as such were entitled to be designated, according to the ordinary rules of classification, by the same names. He also observed that other shells were so nearly allied to living forms that it was difficult not to suspect that they had been connected by a common bond of descent. He therefore proposed that the element of time should enter into the definition of a species, and that it should run thus: "A species consists of individuals all resembling each other, and reproducing their like by generation, SO LONG AS THE SURROUNDING CONDITIONS DO NOT UNDERGO CHANGES SUFFICIENT TO CAUSE THEIR HABITS, CHARACTERS, AND FORMS TO VARY." He came at last to the conclusion that none of the animals and plants now existing were primordial creations, but were all derived from pre-existing forms, which, after they may have gone on for indefinite ages reproducing their like, had at length, by the influence of alterations in climate and in the animate world been made to vary gradually, and adapt themselves to new circumstances, some of them deviating in the course of ages so far from their original type as to have claims to be regarded as new species.
In support of these views, he referred to wild and cultivated plants and to wild and domesticated animals, pointing out how their colour, form, structure, physiological attributes and even instincts were gradually modified by exposure to new soils and climates, new enemies, modes of subsistence, and kinds of food.