HKB
(* "Zeitschrift der Deutschen Geologischen Gesellschaft" volume 7 1855 page 497 etc.)
As an example of what may possibly have constituted a more southern fauna in the valley of the Thames, I may allude to the fossil remains found in the fluviatile alluvium of Gray's Thurrock, in Essex, situated on the left bank of the river, 21 miles below London. The strata of brick-earth, loam, and gravel exposed to view in artificial excavations in that spot, are precisely such as would be formed by the silting up of an old river channel. Among the mammalia are Elephas antiquus, Rhinoceros leptorhinus (R. megarhinus, Christol), Hippopotamus major, species of horse, bear, ox, stag, etc., and, among the accompanying shells, Cyrena fluminalis, which is extremely abundant, instead of being scarce, as at Abbeville. It is associated with Unio littoralis also in great numbers and with both valves united. This conspicuous freshwater mussel is no longer an inhabitant of the British Isles, but still lives in the Seine, and is still more abundant in the Loire. Another freshwater univalve (Paludina marginata, Michaud), not British, but common in the south of France, likewise occurs, and a peculiar variety of Cyclas amnica, which by some naturalists has been regarded as a distinct species. With these, moreover, is found a peculiar variety of Valvata piscinalis.
If we consult Dr. Von Schrenck's account of the living mammalia of Mongolia, lying between latitude 45 and 55 degrees north, we learn that, in that part of North-Eastern Asia recently annexed to the Russian empire, no less than thirty-four out of fifty-eight living quadrupeds are identical with European species, while some of those which do not extend their range to Europe are arctic, others tropical forms. The Bengal tiger ranges northwards occasionally to latitude 52 degrees north, where he chiefly subsists on the flesh of the reindeer, and the same tiger abounds in latitude 48 degrees, to which the small tailless hare or pika, a polar resident, sometimes wanders southwards.*
(* Mammalia of Amoorland, "Natural History Review" volume 1 1861 page 12.)
We may readily conceive that the countries now drained by the Thames, the Somme, and the Seine, were, in the Pleistocene period, on the borders of two distinct zoological provinces, one lying to the north, the other to the south, in which case many species belonging to each fauna endowed with migratory habits, like the living musk-ox or the Bengal tiger, may have been ready to take advantage of any, even the slightest, change in their favour to invade the neighbouring province, whether in the summer or winter months, or permanently for a series of years, or centuries. The Elephas antiquus and its associated Rhinoceros leptorhinus may have preceded the mammoth and tichorhine rhinoceros in the valley of the Thames, or both may have alternately prevailed in the same area in the Pleistocene period.