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Fig. 91), concealed in the socket. From the fact of this molar not having been cut, we learn that the individual was young, and had not shed its first teeth.
The reader will observe that all these extinct quadrupeds of Australia belong to the marsupial family, or, in other words, that they are referable to the same peculiar type of organisation which now distinguishes the Australian mammalia from those of other parts of the globe. This fact is one of many pointing to a general law deducible from the fossil vertebrate and invertebrate animals of times immediately antecedent to our own, namely, that the present geographical distribution of organic forms dates back to a period anterior to the origin of existing species; in other words, the limitation of particular genera or families of quadrupeds, mollusca, etc., to certain existing provinces of land and sea, began before the larger part of the species now contemporary with man had been introduced into the earth.
Professor Owen, in his excellent “History of British Fossil Mammals,” has called attention to this law, remarking that the fossil quadrupeds of Europe and Asia differ from those of Australia or South America. We do not find, for example, in the Europæo-Asiatic province fossil kangaroos, or armadillos, but the elephant, rhinoceros, horse, bear, hyæna, beaver, hare, mole, and others, which still characterise the same continent.
In like manner, in the Pampas of South America the skeletons of Megatherium, Megalonyx, Glyptodon, Mylodon, Toxodon, Macrauchenia, and other extinct forms, are analogous to the living sloth, armadillo, cavy, capybara, and llama. The fossil quadrumana, also associated with some of these forms in the Brazilian caves, belong to the Platyrrhine family of monkeys, now peculiar to South America. That the extinct fauna of Buenos Ayres and Brazil was very modern has been shown by its relation to deposits of marine shells, agreeing with those now inhabiting the Atlantic.
The law of geographical relationship above alluded to, between the living vertebrata of every great zoological province and the fossils of the period immediately antecedent, even where the fossil species are extinct, is by no means confined to the mammalia. New Zealand, when first examined by Europeans, was found to contain no indigenous land quadrupeds, no kangaroos, or opossums, like Australia; but a wingless bird abounded there, the smallest living representative of the ostrich family, called the Kiwi by the natives (Apteryx). In the fossils of the Post-pliocene period in this same island, there is the like absence of kangaroos, opossums, wombats, and the rest; but in their place a prodigious number of well-preserved specimens of gigantic birds of the struthious order, called by Owen Dinornis and Palapteryx, which are entombed in superficial deposits. These genera comprehended many species, some of which were four, some seven, others nine, and others eleven feet in height! It seems doubtful whether any contemporary mammalia shared the land with this population of gigantic feathered bipeds.
Mr. Darwin, when describing the recent and fossil mammalia of South America, has dwelt much on the wonderful relationship of the extinct to the living types in that part of the world, inferring from such geographical phenomena that the existing species are all related to the extinct ones which preceded them by a bond of common descent.