HKB
that in South America a native horse should have lived and disappeared, to be succeeded in after-ages by the countless herds descended from the few introduced with the Spanish colonists!
The existence in South America of a fossil horse, of the mastodon, possibly of an elephant, [4] and of a hollow-horned ruminant, discovered by MM. Lund and Clausen in the caves of Brazil, are highly interesting facts with respect to the geographical distribution of animals. At the present time, if we divide America, not by the Isthmus of Panama, but by the southern part of Mexico [5] in lat. 20 degs., where the great table-land presents an obstacle to the migration of species, by affecting the climate, and by forming, with the exception of some valleys and of a fringe of low land on the coast, a broad barrier; we shall then have the two zoological provinces of North and South America strongly contrasted with each other. Some few species alone have passed the barrier, and may be considered as wanderers from the south, such as the puma, opossum, kinkajou, and peccari. South America is characterized by possessing many peculiar gnawers, a family of monkeys, the llama, peccari, tapir, opossums, and, especially, several genera of Edentata, the order which includes the sloths, ant-eaters, and armadilloes. North America, on the other hand, is characterized (putting on one side a few wandering species) by numerous peculiar gnawers, and by four genera (the ox, sheep, goat, and antelope) of hollow-horned ruminants, of which great division South America is not known to possess a single species. Formerly, but within the period when most of the now existing shells were living, North America possessed, besides hollow-horned ruminants, the elephant, mastodon, horse, and three genera of Edentata, namely, the Megatherium, Megalonyx, and Mylodon. Within nearly this same period (as proved by the shells at Bahia Blanca) South America possessed, as we have just seen, a mastodon, horse, hollow-horned ruminant, and the same three genera (as well as several others) of the Edentata. Hence it is evident that North and South America, in having within a late geological period these several genera in common, were much more closely related in the character of their terrestrial inhabitants than they now are. The more I reflect on this case, the more interesting it appears: I know of no other instance where we can almost mark the period and manner of the splitting up of one great region into two well-characterized zoological provinces. The geologist, who is fully impressed with the vast oscillations of level which have affected the earth's crust within late periods, will not fear to speculate on the recent elevation of the Mexican platform, or, more probably, on the recent submergence of land in the West Indian Archipelago, as the cause of the present zoological separation of North and South America. The South American character of the West Indian mammals [6] seems to indicate that this archipelago was formerly united to the southern continent, and that it has subsequently been an area of subsidence.
When America, and especially North America, possessed its elephants, mastodons, horse, and hollow-horned ruminants, it was much more closely related in its zoological characters to the temperate parts of Europe and Asia than it now is. As the remains of these genera are found on both sides of Behring's Straits [7] and on the plains of Siberia, we are led to look to the north-western side of North America as the former point of communication between the Old and so-called New World. And as so many species, both living and extinct, of these same genera inhabit and have inhabited the Old World, it seems most probable that the North American elephants, mastodons, horse, and hollow-horned ruminants migrated, on land since submerged near Behring's Straits, from Siberia into North America, and thence, on land since submerged in the West Indies, into South America, where for a time they mingled with the forms characteristic of that southern continent, and have since become extinct.